Publications Bernhard A. Huber      last update: November 2011

In press Huber, B. A. In press. Pholcidae. In: Biodiversidad de Artrópodos Argentinos, Vol. 3 (eds. L Claps, G Debandi, y SR Juñent). Sociedad Entomológica Argentina. 2011 Huber, B. A. 2011. Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1-509. PDF (high resolution, 206 MB) PDF (low resolution, 36 MB) Huber, B. A. 2011. Phylogeny and classification of Pholcidae (Araneae): an update. Journal of Arachnology 39: 211-222. PDF Huber, B. A., Rheims, C. A. 2011. Diversity and endemism of pholcid spiders in Brazil’s Atlantic Forest, with descriptions of four new species of the Atlantic Forest endemic genus Tupigea (Araneae: Pholcidae). Journal of Natural History 45: 275-301. PDF request 2010 Huber, B. A. 2010. Mating positions and the evolution of asymmetric insect genitalia. Genetica 138: 19-25. PDF request Huber, B. A., Fischer, N., Astrin, J. J. 2010. High level of endemism in Haiti's last remaining forests: a revision of Modisimus (Araneae: Pholcidae) on Hispaniola, using morphology and molecules. Zoological Journal of the Linnean Society 158: 244-299. PDF request Huber, B. A., Lankhorst, S. 2010. Non-insect arthropod types in the ZFMK collection, Bonn (Acari, Araneae, Scorpiones, Pantopoda, Amphipoda). Bonn zoological Bulletin 58: 217-226. PDF Eberhard, W. G., Huber, B. A. 2010. Spider genitalia: precise maneuvers with a numb structure in a complex lock. Pp. 249-284. In: J. L. Leonard & A. Córdoba-Aguilar (eds.) Evolution of primary sexual characters in animals. Oxford University Press, Oxford, U.K. PDF 2009 Huber, B. A. 2009. Life on leaves: leaf-dwelling pholcids of Guinea, with emphasis on Crossopriza cylindrogaster, a spider with inverted resting position, pseudo-eyes, lampshade web, and tetrahedral egg-sac (Araneae: Pholcidae). Journal of Natural History 43: 2491-2523. PDF request Huber, B. A. 2009. Four new generic and 14 new specific synonymies in Pholcidae, and transfer of Pholcoides Roewer to Filistatidae (Araneae). Zootaxa 1970: 64-68; 1977: 68 (erratum). PDF request Huber, B. A., Astrin, J. J. 2009. Increased sampling blurs morphological and molecular species limits: revision of the Hispaniolan endemic spider genus Tainonia (Araneae: Pholcidae). Invertebrate Systematics 23: 281-300. PDF request Huber, B. A., Schütte, A. 2009. Preliminary notes on leaf-dwelling Metagonia spiders (Araneae: Pholcidae) in the Esquinas Rainforest near La Gamba, Costa Rica: leaf preference, mimesis, and web structure. Contributions to Natural History (Bern) 12: 681-697. PDF 2008 Huber, B. A., Fleckenstein, N. 2008. Comb-hairs on the fourth tarsi in pholcid spiders (Araneae, Pholcidae). Journal of Arachnology 36: 232-240. PDF Beatty, J. A, Berry, J. W., Huber, B. A. 2008. The pholcid spiders of Micronesia and Polynesia (Araneae, Pholcidae). Journal of Arachnology 36: 1-25. PDF 2007 Astrin, J. J., Misof, B., Huber, B. A. 2007. The pitfalls of exageration: molecular and morphological evidence suggests Kaliana is a synonym of Mesabolivar (Araneae: Pholcidae). Zootaxa 1646: 17-30. PDF Huber, B. A. 2007. Two new genera of small, six-eyed pholcid spiders from West Africa, and first record of Spermophorides for mainland Africa (Araneae: Pholcidae). Zootaxa 1635: 23-43. PDF Huber, B. A., El Hennawy, H. 2007. On Old World ninetine spiders (Araneae: Pholcidae), with a new genus and species and the first record for Madagascar. Zootaxa 1635: 45-53. PDF Huber, B. A., Sinclair, B., Schmitt, M. 2007. The evolution of asymmetric genitalia in spiders and insects. Biological Reviews 82: 647-698. PDF Machado, É. O., Brescovit, A. D., Candiani, D. F., Huber, B. A. 2007. Three new species of Mesabolivar (Aranea, Pholcidae) from leaf litter in urban environments in the city of São Paulo, São Paulo, Brazil. Iheringia 97: 168-176. PDF Machado, É. O., Yamamoto, F. U., Brescovit, A. D., Huber, B. A. 2007. Three new ground living pholcid species (Araneae: Pholcidae) from Parque Estadual da Cantareira, São Paulo, São Paulo, Brazil. Zootaxa 1582: 27–37. PDF Useta, G., Huber, B. A., Costa, F. G. 2007. Spermathecal morphology and sperm dynamics in the female Schizocosa malitiosa (Araneae: Lycosidae). European Journal of Entomology 104: 777-785. PDF 2006 Astrin J. J., Huber, B. A., Misof B., Kluetsch, C.F.C. 2006. Molecular taxonomy in pholcid spiders (Pholcidae, Araneae): evaluations of species identification methods using CO1 and 16S rRNA. Zoologica Scripta 35: 441-457.  PDF Huber, B. A. 2006. Cryptic female exaggeration: the asymmetric female internal genitalia of Kaliana yuruani (Araneae: Pholcidae). Journal of Morphology 276: 705-712.  PDF Huber, B. A., Wunderlich, J. 2006. Fossil and extant species of the genus Leptopholcus in the Dominican Republic, with the first case of egg-parasitism in pholcid spiders (Araneae: Pholcidae). Journal of Natural History 40: 2341-2360.  PDF Michalik, P., Huber, B. A. 2006. Spermiogenesis in Psilochorus simoni (Berland, 1911) (Pholcidae, Araneae): evidence for considerable within-family variation in sperm structure and development. Zoology 109: 14-25. PDF 2005 Bruvo-Madaric B., Huber, B. A., Steinacher, A., Pass G. 2005. Phylogeny of pholcid spiders (Araneae:Pholcidae): combined analysis using morphology and molecules. Molecular Phylogenetics and Evolution 37: 661-673.  PDF Funke, S., Huber, B. A. 2005. Allometry of genitalia and fighting structures in Linyphia triangularis (Araneae: Linyphiidae). Journal of Arachnology 33: 870-872.  PDF Huber, B. A. 2005a. High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica 155: 1-126.  PDF (21MB) Huber, B. A. 2005b. Sexual selection research on spiders: progress and biases. Biological Reviews 80: 363-385.  PDF Huber, B. A. 2005c. Revision of the genus Spermophora Hentz in Southeast Asia and on the Pacific Islands, with descriptions of three new genera (Araneae: Pholcidae). Zoologische Mededelingen 79-2(4): 61-172.  PDF Huber, B. A. 2005d. Pholcidae. In: Ubick, D., Paquin, P., Cushing, P.E., Roth, V. (eds) Spiders of North America: an identification manual, pp. 194-196.  PDF Huber, B. A. 2005e. The pholcid spiders of Africa (Araneae: Pholcidae): state of knowledge and directions for future research. In: Huber B.A., Sinclair, B.J., Lampe, K.-H. (eds.) African Biodiversity: Molecules, Organisms, Ecosystems, pp. 181-186. Springer Verlag.  PDF Huber, B. A. 2005f. Evidence for directional selection on male abdomen size in Mecolaesthus longissimus (Araneae, Pholcidae). Journal of Arachnology 33: 573-581. PDF Huber, B. A. 2005g. Revision and cladistic analysis of the spider genus Carapoia González-Sponga (Araneae: Pholcidae), with descriptions of new species from Brazil’s Atlantic forest. Invertebrate Systematics 19: 541-556.  PDF Huber, B. A., Benjamin, S. 2005. The pholcid spiders from Sri Lanka: redescription of Pholcus ceylonicus and description of Wanniyala new genus (Araneae: Pholcidae). Journal of Natural History 39(37): 3305-3319. PDF Huber, B. A., Brescovit, A. D. & Rheims, C. A. 2005. Exaggerated female genitalia in two new spider species (Araneae: Pholcidae), with comments on genital evolution by female choice versus antagonistic coevolution. Insect Systematics and Evolution 36: 285-292.  PDF Huber, B. A., Pérez G., A., Baptista, R.L.C. 2005. Leptopholcus (Araneae: Pholcidae) in continental America: rare relicts in low precipitation areas. Bonner zoologische Beiträge 53(1/2): 99-107.  PDF (5MB) Huber, B. A., Rheims, C. A. & Brescovit, A. D. 2005a. Speciation without changes in genital shape: a case study on Brazilian pholcid spiders (Araneae: Pholcidae). Zoologischer Anzeiger 243: 273-279. PDF Huber, B. A., Rheims, C. A. & Brescovit, A. D. 2005b. Two new species of litter-dwelling Metagonia spiders (Araneae, Pholcidae) document both rapid and slow genital evolution. Acta Zoologica (Stockholm): 86: 33-40.  PDF 2004 Huber, B. A. 2004a. Evidence for functional segregation in the directionally asymmetric male genitalia of the spider Metagonia mariguitarensis (Gonzalez-Sponga) (Pholcidae: Araneae). Journal of Zoology, London 262: 317-326.  PDF Huber, B. A. 2004b. Evolutionary transformation from muscular to hydraulic movements in spider (Arachnida, Araneae) genitalia: a study based on histological serial sections. Journal of Morphology 261: 364-376.  PDF Huber, B. A. 2004c. The significance of copulatory structures in spider systematics. In: Schult, J. (ed.) Studien zur Theorie der Biologie, Band 5, Biosemiotik - Praktische Anwendung und Konsequenzen fuer die Einzeldisziplinen, VWB-Verlag fuer Wissenschaft und Bildung, Berlin.  PDF Huber, B. A., Hopf, S. 2004. Color dimorphism in adults and juveniles of Buitinga safura (Araneae, Pholcidae). Journal of Arachnology 32: 336-340.  PDF 2003 Huber, B. A. 2003a. Cladistic analysis of Malagasy pholcid spiders reveals generic level endemism: Revision of Zatavua n. gen. and Paramicromerys Millot (Pholcidae, Araneae). Zoological Journal of the Linnean Society 137: 261-318. PDF Huber, B. A. 2003b. High species diversity in one of the dominant groups of spiders in East African montane forests (Araneae: Pholcidae: Buitinga n. gen., Spermophora Hentz). Zoological Journal of the Linnean Society 137: 555-619. PDF (5MB) Huber, B. A. 2003c. Southern African pholcid spiders: revision and cladistic analysis of Quamtana gen. nov. and Spermophora Hentz (Araneae: Pholcidae), with notes on male-female covariation. Zoological Journal of the Linnean Society 139: 477-527.  PDF Huber, B. A. 2003d. On the terminology of polytomies. Cladistics 19: 273.  PDF (proofs) Huber, B. A. 2003e. Rapid evolution and species-specificity of arthropod genitalia: fact or artifact? Organisms Diversity & Evolution 3: 63-71.  PDF Huber, B. A., Brescovit, A. D. 2003. Ibotyporanga Mello-Leitão: tropical spiders in Brazilian semi-arid habitats (Araneae: Pholcidae). Insect Systematics and Evolution 34: 15-20. PDF 2002 Huber, B. A. 2002a. Functional morphology of the genitalia in the spider Spermophora senoculata (Pholcidae, Araneae). Zoologischer Anzeiger 241: 105-116.  PDF (5MB) Huber, B. A. 2002b. Ninetis russellsmithi n. sp., an unusual new pholcid spider species from Malawi (Araneae: Pholcidae). Journal of Insect Science 2.4: 1-3.  PDF 2001 Huber, B. A. 2001. The pholcids of Australia (Araneae; Pholcidae): taxonomy, biogeography, and relationships. Bulletin of the American Museum of Natural History 260: 1-144. PDF (20MB) Huber, B. A., Pérez González, A. 2001a. A new genus of pholcid spiders (Araneae: Pholcidae) endemic to Western Cuba, with a case of female genitalic dimorphism. American Museum Novitates 3329: 1-23. PDF (11MB) Huber, B. A., Pérez González, A. 2001b. Female genital dimorphism in a spider (Araneae: Pholcidae). Journal of Zoology, London 255: 301-304. PDF Huber, B. A., van Harten, A. 2001. Ninetis subtilissima Simon, 1890 (Araneae: Pholcidae): Redescription and SEM ultrastructure. American Museum Novitates 3336: 1-7.  PDF Huber, B. A., Zhu, M.-S. 2001. On Taczanowski’s pholcids, with three new synonymies in Pholcidae (Araneae). Bulletin of the British arachnological Society 12(3): 151-152. PDF 2000 Huber, B. A. 2000. New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1-348. PDF (8MB) 1999 Eberhard, W. G., Huber, B. A., Rodriguez, R. L. 1999. Don’t forget the biology: a reply to Green. Evolution 53(5): 1624-1627. PDF Huber, B. A. 1999. Sexual selection in pholcid spiders (Araneae, Pholcidae): artful chelicerae and forceful genitalia. Journal of Arachnology 27(1): 135-141.  PDF Huber, B. A., Deeleman-Reinhold, C. L., Pérez González, A. 1999. The spider genus Crossopriza (Araneae, Pholcidae) in the New World. American Museum Novitates 3262: 1-10. PDF Pérez González, A., Huber, B. A. 1999. Metagonia debrasi n. sp., the first species of the genus Metagonia Simon in Cuba (Pholcidae, Araneae). Revue Arachnologique 13(4): 69-72.  PDF 1998 Eberhard, W. G., Huber, B. A. 1998a. Courtship, copulation, and sperm transfer in Leucauge mariana (Araneae, Tetragnathidae) with implications for higher classification. Journal of Arachnology 26(3): 342-368.  PDF Eberhard, W. G., Huber, B. A. 1998b. Possible links between embryology, lack of innervation, and the evolution of male genitalia in spiders. Bulletin of the British arachnological Society 11(2): 73-80.  PDF Eberhard, W. G., Huber, B. A. 1998c. Copulation and sperm transfer in Archisepsis flies (Diptera, Sepsidae) and the evolution of their intromittent genitalia. Studia Dipterologica 5(2): 217-248. Eberhard, W. G., Huber, B. A., Rodriguez, R. L., Briceño, D., Salas, I., Rodriguez, V. 1998. One size fits all? Relationships between the size of genitalia and other body parts in 20 species of insects and spiders. Evolution 52(2): 415-431.  PDF Huber, B. A. 1998a. On the "valve" in the genitalia of female pholcids (Pholcidae, Araneae). Bulletin of the British arachnological Society 11(2): 41-48.  PDF Huber, B. A. 1998b. Notes on the neotropical spider genus Modisimus (Pholcidae, Araneae), with descriptions of thirteen new species from Costa Rica and neighboring countries. Journal of Arachnology 26: 19-60.  PDF (4MB) Huber, B. A. 1998c. Report of some pholcid spiders collected in Guatemala and Honduras (Araneae, Pholcidae). Revue Suisse de Zoologie 105(1): 49-80. PDF Huber, B. A. 1998d. Genital mechanics in some neotropical pholcid spiders (Araneae: Pholcidae), with implications for systematics. Journal of Zoology, London 244: 587-599. PDF Huber, B. A. 1998e. Spider reproductive behaviour: a review of Gerhardt’s work from 1911-1933, with implications for sexual selection. Bulletin of the British arachnological Society 11(3): 81-91.  PDF Huber, B. A., Pérez González, A. 1998. Leptopholcus delicatulus (Araneae, Pholcidae) is a valid name. Journal of Arachnology 26: 251-256.  PDF 1997 Huber, B. A. 1997a. Evidence for gustatorial courtship in a haplogyne spider (Hedypsilus culicinus: Pholcidae: Araneae). Netherlands Journal of Zoology 47(1): 95-98.  PDF Huber, B. A. 1997b. On American ‘Micromerys’ and Metagonia (Araneae, Pholcidae), with notes on natural history and genital mechanics. Zoologica Scripta 25(4): 341-363.  PDF Huber, B. A. 1997c. Redescriptions of Eugène Simon’s neotropical pholcids (Araneae, Pholcidae). Zoosystema 19(4): 573-612.  PDF Huber, B. A. 1997d. The pholcid spiders of Costa Rica (Aranae: Pholcidae). Revista de Biologia Tropical 45(4): 1583-1634.  PDF Huber, B. A., Eberhard, W. G. 1997. Courtship, copulation, and genital mechanics in Physocyclus globosus (Araneae, Pholcidae). Canadian Journal of Zoology 74: 905-918.  PDF Huber, B. A., Senglet, A. 1997. Copulation with contralateral insertion in entelegyne spiders (Araneae: Entelegynae: Tetragnathidae). Netherlands Journal of Zoology 47(1): 99-102.  PDF 1996 Huber, B. A. 1996a. Genitalia, fluctuating asymmetry, and patterns of sexual selection in Physocyclus globosus (Araneae: Pholcidae). Revue Suisse de Zoologie, suppl. 1996: 289-294. PDF Huber, B. A. 1996b. On the distinction between Modisimus and Hedypsilus (Araneae, Pholcidae), with notes on behavior and natural history. Zoologica Scripta 25(3): 233-240.  PDF 1995 Huber, B. A. 1995a. Genital morphology and copulatory mechanics in Anyphaena accentuata (Anyphaenidae) and Clubiona pallidula (Clubionidae: Araneae). Journal of Zoology, London 235: 689-702.  PDF Huber, B. A. 1995b. The retrolateral tibial apophysis in spiders - shaped by sexual selection? Zoological Journal of the Linnean Society 113: 151-163. PDF Huber, B. A. 1995c. Copulatory mechanism in Holocnemus pluchei and Pholcus opilionoides, with notes on male cheliceral apophyses and stridulatory organs in Pholcidae (Araneae). Acta Zoologica, Stockholm 76(4): 291-300.  PDF Uhl, G., Huber, B. A., Rose, W. 1995. Male pedipalp morphology and copulatory mechanism in Pholcus phalangioides (Fuesslin, 1775) (Araneae, Pholcidae). Bulletin of the British arachnological Society 10(1): 1-9.  PDF 1994 Huber, B. A. 1994a. Spermophore morphology reveals a new synapomorphy of Oecobius and Uroctea (Araneae, Oecobiidae). Journal of Arachnology 22: 73-74.  PDF Huber, B. A. 1994b. Copulatory mechanics in the funnel-web spiders Histopona torpida and Textrix denticulata (Agelenidae, Araneae). Acta Zoologica, Stockholm 75(4): 379-384. PDF Huber, B. A. 1994c. Funktion und Evolution komplexer Kopulationsorgane. Eine Analyse von Schnittserien durch in copula schockfixierte Spinnen. Mitt. dtsch. Ges. allg. angew. Entomol. 9: 247-250. Huber, B. A. 1994d. Genital bulb muscles in entelegyne spiders. Journal of Arachnology 22: 75-76.  PDF Huber, B. A. 1994e. Genital morphology, copulatory mechanism and reproductive biology in Psilochorus simoni (Berland, 1911) (Pholcidae; Araneae). Netherlands Journal of Zoology 44(1-2): 85-99.  PDF 1993 Huber, B. A. 1993a. Genital mechanics and sexual selection in the spider Nesticus cellulanus (Araneae: Nesticidae). Canadian Journal of Zoology 71: 2437-2447.  PDF Huber, B. A. 1993b. Female choice and spider genitalia. Boll. Accad. Gioenia Sci. Nat. (Catania) 26: 209-214.  PDF Huber K. C., Haider T. S., Müller M. W., Huber B. A., Schweyen R. J., Barth F. G. 1993. DNA sequence data indicates the polyphyly of the family Ctenidae (Araneae). Journal of Arachnology 21: 194-201. PDF 1992 Huber, B. A. 1992. Frontal heart and arterial system in the head of isopoda. Crustaceana 63(1): 57-69. PDF